Ray Romano
Manny (voice)

John Leguizamo
Sid (voice)

Denis Leary
Diego (voice)

Seann William Scott
Crash (voice)

Josh Peck
Eddie (voice)

Queen Latifah
Ellie (voice)

Jennifer Lopez
Shira (voice)

Peter Dinklage
Captain Gutt (voice)

Wanda Sykes
Granny (voice)

Keke Palmer
Peaches (voice)

Josh Gad
Louis (voice)

Nick Frost
Flynn (voice)

Aziz Ansari
Squint (voice)

Alain Chabat
Silas (voice)

Kunal Nayyar
Gupta (voice)

Drake
Ethan (voice)

Nicki Minaj
Steffie (voice)

Rebel Wilson
Raz (voice)

Alan Tudyk
Milton / Hunky Siren (voice)

Ester Dean
Female Siren (voice)

Eddie 'Piolin' Sotelo
Uncle Fungus (voice)

Joy Behar
Eunice (voice)

Heather Morris
Katie (voice)

Chris Wedge
Scrat (voice)

Karen Disher
Scratte (voice)

Marieve Herington
Additional Voices (voice)

Allegra Leguizamo
Additional Voices (voice)

Jason Fricchione
Dumb Mammoth / Various (voice)

Ben Gleib
Marshall (voice)

Simon Pegg
Buck (voice)

Alexandra Romano
Meghan (voice)

Patrick Stewart
Ariscratle (voice)

Brian Scott McFadden
Additional Voices (voice)

Edita Brychta
Additional Voices (voice)

Dennis T. Carnegie
Additional Voices (voice)

Aimée Castle
Additional Voices (voice)

Jim Conroy
Additional Voices (voice)

Jason Harris
Additional Voices (voice)

Sondra James
Additional Voices (voice)

Alexa Kahn
Additional Voices (voice)

Bill Lobley
Additional Voices (voice)

Chris Phillips
Additional Voices (voice)

Tara Sands
Additional Voices (voice)

Keith Silverstein
Additional Voices (voice)

Vanessa Lemonides
Additional Voices (voice)

Danny Flaherty
Additional Voices (voice)

Selenis Leyva
Additional Voices (voice)

Ron McClary
Additional Voices (voice)

Peter Pamela Rose
Additional Voices (voice)

Gabriel Rush
Additional Voices (voice)

Randy Thom
Additonal Voices (voice)

Annie Maginnis Tippe
Additional Voices (voice)

Corwin C. Tuggles
Additional Voices (voice)

Tashiana Washington
Additional Voices (voice)